Examples

Example workflows for using glycowork in your work

On this page, there are different toy examples demonstrating how you can apply glycowork to your projects! Here is a summary of the different available case studies:

1. Investigation of N-linked and O-linked glycosylation in fish
2. Glycan binding specificities of lectins
3. Glycan classification using machine learning models
4. Constructing and exploring biosynthetic networks
5. Deep Learning Code Snippets

Example 1: Investigation of N-linked and O-linked glycosylations in fish

Suppose that after investigating glycans from plants as presented in the core module, you are now wondering how glycans from the Actinopterygii class (ray-finned fish) look like. To satisfy your curiosity, let’s start with importing the df_species dataset from glycowork.glycan_data.loader and putting a filter for animal glycans.

df_animal = df_species[df_species.Class == 'Actinopterygii'].reset_index(drop = True)

	Species	Genus	Family	Order	Class	Phylum	Kingdom	Domain	ref
Gal(?1-?)[Gal(?1-?)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Acipenser_brevirostrum	Acipenser	Acipenseridae	Acipenseriformes	Actinopterygii	Chordata	Animalia	Eukarya	https://www.frontiersin.org/articles/10.3389/fmolb.2021.778383/full
Gal(?1-?)[Gal(?1-?)]Gal(b1-4)GlcNAc(b1-2)[Gal(?1-?)[Gal(?1-?)]Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Acipenser_brevirostrum	Acipenser	Acipenseridae	Acipenseriformes	Actinopterygii	Chordata	Animalia	Eukarya	https://www.frontiersin.org/articles/10.3389/fmolb.2021.778383/full
Gal(?1-?)[Gal(?1-?)]Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Acipenser_oxyrinchus	Acipenser	Acipenseridae	Acipenseriformes	Actinopterygii	Chordata	Animalia	Eukarya	https://www.frontiersin.org/articles/10.3389/fmolb.2021.778383/full
Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Acipenser_oxyrinchus	Acipenser	Acipenseridae	Acipenseriformes	Actinopterygii	Chordata	Animalia	Eukarya	https://www.frontiersin.org/articles/10.3389/fmolb.2021.778383/full
Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Acipenser_oxyrinchus	Acipenser	Acipenseridae	Acipenseriformes	Actinopterygii	Chordata	Animalia	Eukarya	https://www.frontiersin.org/articles/10.3389/fmolb.2021.778383/full

To have a better overview of the glycans similarities in fish, you can plot these data using plot_embeddings from glycowork.motif.analysis. The best option here, to visualize the data, is to color the dots by taxonomic family.

plot_embeddings(df_animal.target.values.tolist(), label_list = df_animal.Family.values.tolist())

Interesting! First, the graph shows a large Salmonidae cluster in addition to maybe two elongated clusters of various families, with a large set of Cyprinidae glycans.

Now, let’s focus on further analyses. We will first convert our data into a count table using presence_to_matrix from the glycowork.motif.processing module and then visualize the results. A heatmap can be generated using make_heatmap from glycowork.motif.analysis, and the yticklabels = True option allows us to display all the labels.

df_map_animal = presence_to_matrix(df_animal, label_col_name = 'Family')
make_heatmap(df_map_animal, mode = 'motif', feature_set = ['known', 'exhaustive'], datatype = 'presence', yticklabels = True,
            estimate_speedup = True)

The first thing that catches the eye is the presence of a bunch of glycan motifs in Mugilidae to Cichlidae that are absent in the other classes. One hypothesis for this is that these families mainly contain glycosphingolipids in our database, while the other families focus on N-linked glycans (and Salmonidae + Cyprinidae have both as well-investigated families).

To visualize which dots from the embedding plot correspond to glycosphingolipids, we can focus on Glc1Cer-containing glycans. We can map them again but with the additional option shape_feature. This parameter allows you to transform dots into crosses if the corresponding glycan contains the specified monosaccharide or linkage.

plot_embeddings(df_animal.target.values.tolist(), label_list = df_animal.Family.values.tolist(), shape_feature = "Glc1Cer")

As you can see, in df_species, Glc1Cer-containing glycans (crosses) form a nice cluster (in the class Actinopterygii). In this class, they even constitute more isolated clusters, presumably corresponding to certain taxonomic groups.

We can also use glycowork to take a look at the overall monosaccharide distribution in animal glycans. The characterize_monosaccharide from glycowork.motif.analysis will be our best ally to learn what other monosaccharides, for instance, Gal variants are often connected to.

characterize_monosaccharide('Gal', rank = 'Kingdom', focus = 'Animalia', modifications = True)

Running this analysis, it appears that Gal is found connected to GlcNAc as well as various other monosaccharides and that it does display several variants, including sulfurylated galactose. This is a completely different profile than we obtained for xylose in plant glycans!

Example 2: Glycan binding specificities of lectins

For this second example, we will focus on a specific set of proteins, called lectins, that bind glycans in a sequence-specific manner. One example for lectins can be found in the hemagglutinin protein of influenza virus. Influenza viruses are well known as the biological agents responsible for the seasonal flu in humans. In general, viruses penetrate inside host cells thanks to a mechanism involving contacts between proteins and glycans. Surface proteins surrounding viruses and eukaryotic cells interact together and allow the entry of viral particles inside the targeted cells. However, different influenza strains may recognize different glycosylations on proteins more or less efficiently.

To measure these glycan binding specificities, one can add virus particles to glycan arrays presenting immobilized glycans. Such a protocol allows screening for specific binding. From such data, we can then ask what are the glycans that are the most efficiently recognized by a given influenza virus?

Fortunately, glycowork can be really helpful to answer such a question! The binding specificity of 1,392 lectins for a vast range of glycans is available in the glycan_binding dataset. Let’s start by importing these data.

	3-Anhydro-Gal(a1-3)Gal(b1-4)3-Anhydro-Gal(a1-3)Gal4S	3-Anhydro-Gal(a1-3)Gal4S(b1-4)3-Anhydro-Gal(a1-3)Gal4S	3-Anhydro-Gal(a1-3)Gal4S(b1-4)3-Anhydro-Gal(a1-3)Gal4S(b1-4)3-Anhydro-Gal(a1-3)Gal4S	3-Anhydro-Gal(a1-3)Gal4S(b1-4)3-Anhydro-Gal(a1-3)Gal4S(b1-4)3-Anhydro-Gal(a1-3)Gal4S(b1-4)3-Anhydro-Gal(a1-3)Gal4S	3-Anhydro-Gal(a1-3)Gal4S(b1-4)3-Anhydro-Gal2S(a1-3)Gal4S(b1-4)3-Anhydro-Gal(a1-3)Gal4S	3dGal(b1-3)[Fuc(a1-4)]Glc	3dGal(b1-4)Glc	4d8dNeu5Ac(a2-3)Gal(b1-4)Glc	4dNeu5Ac(a2-3)Gal(b1-4)Glc	7dNeu5Ac(a2-3)Gal(b1-4)Glc	8dNeu5Ac(a2-3)Gal(b1-4)Glc	9dNeu5Ac(a2-3)Gal(b1-4)Glc	Ara(a1-5)Ara(a1-5)Ara(a1-5)Ara(a1-5)Ara(a1-5)Ara	Ara(a1-5)Ara(a1-5)Ara(a1-5)Ara(a1-5)Ara(a1-5)Ara(a1-5)Ara	Fuc(a1-2)Gal	Fuc(a1-2)Gal(b1-3)GalNAc	Fuc(a1-2)Gal(b1-3)GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-3)GalNAc(b1-3)Gal	Fuc(a1-2)Gal(b1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GalNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GalNAc(b1-4)[Neu5Ac(a2-3)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GlcNAc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)GalNAc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-2)Gal(b1-3)GlcNAc(b1-6)]GalNAc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-3)]Gal(b1-4)GlcNAc(b1-6)[Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)GlcNAc6S	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-6)[Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)[Fuc(a1-2)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-4)Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-4)Glc6S	Fuc(a1-2)Gal(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)GalNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-6)]GalNAc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-6)GalNAc	Fuc(a1-2)Gal(b1-4)GlcNAc6S	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GalNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GalNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GalNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)GalNAc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)[Fuc(a1-2)]Gal(b1-4)Glc	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-2)Gal3S(b1-4)GlcNAc	Fuc(a1-2)Gal6S(b1-3)GlcNAc	Fuc(a1-2)Gal6S(b1-3)GlcNAc6S	Fuc(a1-2)Gal6S(b1-4)Glc	Fuc(a1-2)Gal6S(b1-4)Glc6S	Fuc(a1-2)Gal6S(b1-4)GlcNAc	Fuc(a1-2)GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GalNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc(b1-3)GalNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-3)GlcNAc(b1-6)GalNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)Glc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-3)GalNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-3)[Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-6)]GalNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)GlcNAc(b1-6)GalNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)[Fuc(a1-3)]Glc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-2)[Gal(a1-3)]Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)GalNAc	Fuc(a1-2)[Gal(a1-4)]Gal(b1-4)GalNAc	Fuc(a1-2)[Gal(a1-4)]Gal(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc(b1-3)GalNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)GlcNAc(b1-6)GalNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)Glc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-3)GalNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-3)[Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-6)]GalNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)GlcNAc(b1-6)GalNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-2)[GalNAc(a1-3)]Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)GalNAc	Fuc(a1-2)[GalNAc(a1-4)]Gal(b1-4)GlcNAc	Fuc(a1-2)[GalNAc(b1-3)]Gal	Fuc(a1-2)[Neu5Ac(bond)]Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-3)GlcNAc	Fuc(a1-3)GlcNAc6S	Fuc(a1-3)[3dGal(b1-4)]Glc	Fuc(a1-3)[3dGal(b1-4)]GlcNAc(b1-3)Gal	Fuc(a1-3)[Gal(b1-4)]Glc	Fuc(a1-3)[Gal(b1-4)]Glc6S	Fuc(a1-3)[Gal(b1-4)]GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-2)Man	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-2)Man(a1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-2)Man(a1-6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]Glc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)GalNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)[Fuc(a1-2)]Gal(b1-4)Glc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-4)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-4)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)GalNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)[Gal(b1-3)GlcNAc(b1-3)]Gal(b1-4)Glc	Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)[Neu5Ac(a2-3)Gal(b1-3)]GalNAc	Fuc(a1-3)[Gal(b1-4)]GlcNAc6S	Fuc(a1-3)[Gal3S(b1-4)]Glc	Fuc(a1-3)[Gal3S(b1-4)]Glc6S	Fuc(a1-3)[Gal3S(b1-4)]GlcNAc	Fuc(a1-3)[Gal3S(b1-4)]GlcNAc(b1-3)Gal	Fuc(a1-3)[Gal3S(b1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-3)[Gal3S(b1-4)]GlcNAc6S	Fuc(a1-3)[Gal3S(b1-4)]GlcNAc6S(b1-3)Gal(b1-4)Glc	Fuc(a1-3)[Gal6S(b1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Fuc(a1-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc	Fuc(a1-3)[GalNAc(b1-4)]GlcNAc	Fuc(a1-3)[GalNAc(b1-4)]GlcNAc6S	Fuc(a1-3)[GalNAc3S(b1-4)]GlcNAc	Fuc(a1-3)[Neu5Ac(a2-3)]Gal(b1-4)GlcNAc(b1-3)GalNAc	Fuc(a1-4)GlcNAc	Fuc(a1-6)GlcNAc	Fuc(b1-3)GlcNAc	Gal(a1-2)Gal	Gal(a1-3)Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)[Gal(a1-3)Fuc(a1-2)Gal(b1-4)GlcNAc(b1-6)]GalNAc	Gal(a1-3)Gal	Gal(a1-3)Gal(b1-3)Glc	Gal(a1-3)Gal(b1-3)GlcNAc	Gal(a1-3)Gal(b1-3)GlcNAc(b1-2)Man	Gal(a1-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(a1-3)Gal(b1-3)GlcNAc(b1-3)GalNAc	Gal(a1-3)Gal(b1-3)GlcNAc(b1-6)GalNAc	Gal(a1-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(a1-3)Gal(b1-4)Gal(a1-3)Gal	Gal(a1-3)Gal(b1-4)Glc	Gal(a1-3)Gal(b1-4)GlcNAc	Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man	Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(a1-3)Gal(b1-4)GlcNAc(b1-3)GalNAc	Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)GalNAc	Gal(a1-3)Gal(b1-4)[Fuc(a1-3)]GalNAc	Gal(a1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Gal(a1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(a1-3)GalNAc	Gal(a1-3)GlcNAc	Gal(a1-3)[Gal(a1-4)]Gal(b1-4)GlcNAc	Gal(a1-4)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(a1-4)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(a1-4)Gal(b1-4)Gal	Gal(a1-4)Gal(b1-4)GalNAc	Gal(a1-4)Gal(b1-4)Glc	Gal(a1-4)Gal(b1-4)GlcNAc	Gal(a1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(a1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(a1-4)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(a1-4)Glc	Gal(a1-4)GlcNAc	Gal(a1-6)Glc	Gal(b1-2)Gal	Gal(b1-3)Gal	Gal(b1-3)Gal(b1-4)GlcNAc	Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-3)GalNAc	Gal(b1-3)GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)Glc	Gal(b1-3)GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc	Gal(b1-3)GalNAc(b1-3)Gal	Gal(b1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc	Gal(b1-3)GalNAc(b1-3)Gal(b1-4)Glc	Gal(b1-3)GalNAc(b1-3)Gal(b1-4)GlcNAc	Gal(b1-3)GalNAc(b1-4)Gal(b1-4)Glc	Gal(b1-3)GalNAc(b1-4)Gal3S(b1-4)Glc	Gal(b1-3)GalNAc(b1-4)[Neu5Ac(a2-3)]Gal(b1-4)Glc	Gal(b1-3)GalNAc(b1-4)[Neu5Gc(a2-3)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc	Gal(b1-3)GlcNAc(a1-3)Gal(b1-4)GlcNAc	Gal(b1-3)GlcNAc(a1-6)Gal(b1-4)GlcNAc	Gal(b1-3)GlcNAc(b1-2)Man	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-3)GlcNAc(b1-2)[Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-3)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-3)GlcNAc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)GlcNAc(b1-3)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)[Gal(b1-3)GlcNAc(b1-3)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)GalNAc	Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)]GlcNAc(b1-6)[Gal(b1-4)]GlcNAc(b1-3)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-3)]Gal(b1-4)GlcNAc(b1-6)[Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)GlcNAc(b1-3)]Gal(b1-4)Glc	Gal(b1-3)GlcNAc(b1-6)Gal(b1-4)GlcNAc	Gal(b1-3)GlcNAc(b1-6)GalNAc	Gal(b1-3)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Gal(b1-3)GlcNAc6S	Gal(b1-3)[Fuc(a1-4)]GlcNAc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)GlcNAc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]Glc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)[Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)]Gal(b1-4)Glc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-6)GalNAc	Gal(b1-3)[GlcNAc(b1-6)]GalNAc	Gal(b1-3)[GlcNAc(b1-6)]GlcNAc	Gal(b1-3)[Neu5Ac(a2-6)]GalNAc	Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(b1-4)Gal(b1-4)Glc	Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc	Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(a1-4)Gal(b1-4)Glc	Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-4)Gal(b1-4)Glc	Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-6)Gal(b1-4)Glc	Gal(b1-3)[Neu5Ac(b2-6)]GalNAc	Gal(b1-3)[Neu5Ac(b2-6)]GlcNAc	Gal(b1-4)Gal	Gal(b1-4)GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc	Gal(b1-4)GalNAc(b1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc	Gal(b1-4)Glc	Gal(b1-4)Glc6S	Gal(b1-4)GlcNAc	Gal(b1-4)GlcNAc(a1-6)Gal(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Gal(b1-4)GlcNAc(b1-2)GalNAc	Gal(b1-4)GlcNAc(b1-2)Man	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)][Gal(b1-4)GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Fuc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GalNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)GalNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)GlcNAc(b1-6)]Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Gal(b1-4)GlcNAc(b1-3)GalNAc	Gal(b1-4)GlcNAc(b1-3)[Fuc(a1-3)GlcNAc(b1-6)]Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc	Gal(b1-4)GlcNAc(b1-3)[GlcNAc(b1-6)]Gal(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-6)Gal(b1-4)GlcNAc	Gal(b1-4)GlcNAc(b1-6)Gal(b1-4)GlcNAc(b1-6)GalNAc	Gal(b1-4)GlcNAc(b1-6)GalNAc	Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GlcNAc	Gal(b1-4)GlcNAc(b1-6)[Gal(b1-4)GlcNAc(b1-3)]GalNAc	Gal(b1-4)GlcNAc6P	Gal(b1-6)Gal	Gal(b1-6)GalNAc	Gal3S(b1-3)GalNAc	Gal3S(b1-3)GalNAc(b1-4)Gal3S(b1-4)Glc	Gal3S(b1-3)GlcNAc	Gal3S(b1-3)[Fuc(a1-4)]GalNAc	Gal3S(b1-3)[Fuc(a1-4)]GlcNAc	Gal3S(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Gal3S(b1-4)Glc	Gal3S(b1-4)Glc6S	Gal3S(b1-4)GlcNAc	Gal3S(b1-4)GlcNAc(b1-3)Gal	Gal3S(b1-4)GlcNAc6S	Gal3S(b1-4)[Fuc(a1-3)]Glc	Gal3S(b1-4)[Fuc(a1-3)]Glc6S	Gal3S(b1-4)[Fuc(a1-3)]GlcNAc	Gal3S(b1-4)[Fuc(a1-3)]GlcNAc6S	Gal3S4S(b1-4)GlcNAc	Gal3S6S(b1-4)GlcNAc	Gal3S6S(b1-4)GlcNAc6S	Gal4S(b1-4)GlcNAc	Gal4S6S(b1-4)GlcNAc	Gal6P(b1-4)GlcNAc	Gal6S(b1-3)GlcNAc	Gal6S(b1-3)GlcNAc6S	Gal6S(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Gal6S(b1-4)Glc	Gal6S(b1-4)Glc6S	Gal6S(b1-4)GlcNAc	Gal6S(b1-4)GlcNAc6S	GalNAc(a1-3)Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)[GalNAc(a1-3)Fuc(a1-2)Gal(b1-4)GlcNAc(b1-6)]GalNAc	GalNAc(a1-3)Gal	GalNAc(a1-3)Gal(b1-4)Glc	GalNAc(a1-3)GalNAc	GalNAc(a1-3)GalNAc(a1-3)Gal(a1-4)Gal(b1-4)GlcNAc	GalNAc(a1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc	GalNAc(a1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)GlcNAc	GalNAc(b1-3)Gal	GalNAc(b1-3)Gal(a1-4)Gal(b1-4)GlcNAc	GalNAc(b1-3)Gal(a1-4)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	GalNAc(b1-3)Gal(a1-6)Gal(b1-4)Glc	GalNAc(b1-3)GalNAc	GalNAc(b1-3)GlcNAc	GalNAc(b1-4)Gal(b1-4)Glc	GalNAc(b1-4)Gal3S(b1-4)Glc	GalNAc(b1-4)GlcNAc	GalNAc(b1-4)GlcNAc(b1-2)Man	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GalNAc(b1-4)GlcNAc(b1-3)GalNAc(b1-4)GlcNAc	GalNAc(b1-4)GlcNAc6S	GalNAc(b1-4)[Neu5Ac(a2-6)]Gal(b1-4)Glc	GalNAc(b1-6)GalNAc	GalNAc3S(b1-4)Gal3S(b1-4)Glc	GalNAc3S(b1-4)GlcNAc	GalNAc3S(b1-4)GlcNAc3S	GalNAc3S(b1-4)[Fuc(a1-3)]GlcNAc	GalNAc4S6S(b1-4)GlcNAc	GalNAc6S(b1-4)GlcNAc	Glc(a1-2)Glc	Glc(a1-2)Glc(a1-2)Glc	Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc	Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc	Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc	Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc	Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc	Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc(a1-2)Glc	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc	Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc(a1-3)Glc	Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-4)Glc(a1-6)Glc	Glc(a1-6)Glc	Glc(a1-6)Glc(a1-4)Glc	Glc(a1-6)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-6)Glc(a1-4)Glc(a1-4)Glc(a1-6)Glc(a1-4)Glc(a1-4)Glc	Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc(a1-6)Glc	Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc(b1-2)Glc	Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)[Glc(b1-6)]Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)[Glc(b1-6)]Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)[Glc(b1-6)]Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)Glc(b1-3)[Glc(b1-6)]Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)[Glc(b1-6)]Glc(b1-3)Glc	Glc(b1-3)Glc(b1-3)[Glc(b1-6)]Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-3)Glc(b1-4)Glc	Glc(b1-3)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-3)Glc(b1-4)Glc(b1-6)Glc(b1-4)Glc	Glc(b1-3)[Glc(b1-6)]Glc(a1-3)Glc(b1-3)[Glc(b1-6)]Glc	Glc(b1-3)[Glc(b1-6)]Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc(b1-3)Glc	Glc(b1-4)Glc	Glc(b1-4)Glc(b1-3)Glc	Glc(b1-4)Glc(b1-3)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-3)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-3)Glc(b1-4)Glc(b1-4)Glc(b1-3)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-3)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc(b1-4)Glc	Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc(b1-6)Glc	GlcA(b1-3)Gal	GlcA(b1-3)Gal(b1-3)GlcNAc	GlcA(b1-3)Gal(b1-4)Glc	GlcA(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	GlcA(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	GlcA(b1-3)GlcNAc	GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc	GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc	GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc(b1-4)GlcA(b1-3)GlcNAc	GlcA(b1-4)GlcNAc(a1-4)GlcA(b1-4)2,5-Anhydro-Man	GlcA(b1-4)GlcNAc(a1-4)GlcA(b1-4)GlcNAc(a1-4)GlcA(b1-4)GlcNAc(a1-4)GlcA(b1-4)2,5-Anhydro-Man	GlcA(b1-6)Gal	GlcA3S(b1-3)Gal	GlcA3S(b1-3)Gal(b1-4)Glc	GlcA3S(b1-3)Gal(b1-4)GlcNAc(b1-2)Man	GlcA3S(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	GlcA3S(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	GlcNAc(a1-3)Gal(b1-4)GlcNAc	GlcNAc(a1-4)Gal	GlcNAc(a1-4)Gal(b1-3)GlcNAc	GlcNAc(a1-4)Gal(b1-3)[Fuc(a1-2)]Gal	GlcNAc(a1-4)Gal(b1-4)GalNAc	GlcNAc(a1-4)Gal(b1-4)GlcNAc	GlcNAc(a1-4)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	GlcNAc(a1-4)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	GlcNAc(a1-4)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	GlcNAc(a1-4)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	GlcNAc(a1-6)Gal(b1-4)GlcNAc	GlcNAc(b1-2)Fuc	GlcNAc(b1-2)Gal(b1-3)GalNAc	GlcNAc(b1-2)Gal(b1-3)GlcNAc	GlcNAc(b1-2)Man	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-2)Man(a1-3)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-3)Fuc	GlcNAc(b1-3)Gal	GlcNAc(b1-3)Gal(b1-3)GalNAc	GlcNAc(b1-3)Gal(b1-4)Glc	GlcNAc(b1-3)Gal(b1-4)GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)GalNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)GalNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)[GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)]GalNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)[GlcNAc(b1-3)Gal(b1-3)]GalNAc	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)[GlcNAc(b1-3)]Gal(b1-4)GlcNAc	GlcNAc(b1-3)GalNAc	GlcNAc(b1-3)Man	GlcNAc(b1-3)[GlcNAc(b1-4)][GlcNAc(b1-6)]GlcNAc	GlcNAc(b1-3)[GlcNAc(b1-6)]Gal	GlcNAc(b1-3)[GlcNAc(b1-6)]Gal(b1-4)GlcNAc	GlcNAc(b1-3)[GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)[GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	GlcNAc(b1-3)[GlcNAc(b1-6)]GalNAc	GlcNAc(b1-3)[GlcNAc(b1-6)]GlcNAc	GlcNAc(b1-3)[GlcNAc(b1-6)]GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)Fuc	GlcNAc(b1-4)Gal(b1-4)GlcNAc	GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc	GlcNAc(b1-4)[GlcNAc(b1-6)]GalNAc	GlcNAc(b1-6)Gal(b1-4)GlcNAc	GlcNAc(b1-6)GalNAc	GlcNAc3Prop(b1-4)GlcNAc	GlcNAc6S(b1-3)Gal(b1-4)GlcNAc	HexA(a1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S	HexA(a1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S	HexA(a1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S	HexA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S	HexA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S	HexA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S(b1-4)IdoA(a1-3)GalNAc4S	HexA(a1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S	HexA(a1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S	HexA(a1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S	HexA(a1-4)GlcNAc(a1-4)HexA(a1-4)GlcNAc(a1-4)HexA(a1-4)GlcNAc(a1-4)HexA(a1-4)Man	HexA2S(a1-4)GlcNS6S	HexA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S	HexA6S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S	HexA6S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S(a1-4)IdoA2S(a1-4)GlcNS6S	Kdn(a2-3)Gal(b1-3)GalNAc	Kdn(a2-3)Gal(b1-3)GalNAc(b1-4)[Kdn(a2-3)]Gal(b1-4)Glc	Kdn(a2-3)Gal(b1-3)GlcNAc	Kdn(a2-3)Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Kdn(a2-3)Gal(b1-4)Glc	Kdn(a2-3)Gal(b1-4)GlcNAc	Kdn(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Kdn(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal	Kdn(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Kdn(a2-3)Gal(b1-4)[Rha(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Kdn(a2-3)[GalNAc(b1-4)]Gal(b1-3)GlcNAc	Kdn(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc	Kdn(a2-6)Gal	Kdn(a2-6)Gal(b1-4)GlcNAc	Man(a1-2)Man	Man(a1-2)Man(a1-2)Man(a1-3)Man	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)[Man(a1-3)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-2)Man(a1-6)[Man(a1-3)]Man	Man(a1-2)Man(a1-3)Man	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)[Man(a1-3)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)[Man(a1-2)Man(a1-2)Man(a1-3)]Man	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)[Man(a1-2)Man(a1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-3)[Man(a1-6)]Man(a1-6)[Man(a1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-2)Man(a1-6)[Man(a1-3)]Man(a1-6)[Man(a1-2)Man(a1-2)Man(a1-3)]Man	Man(a1-2)Man(a1-6)[Man(a1-3)]Man(a1-6)[Man(a1-2)Man(a1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-3)Man	Man(a1-3)Man(a1-6)[Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-3)Man(b1-4)GlcNAc	Man(a1-3)[Man(a1-6)]Man	Man(a1-3)[Man(a1-6)]Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-3)[Man(a1-6)]Man(a1-6)[Man(a1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-3)[Man(a1-6)]Man(a1-6)[Man(a1-3)]Man	Man(a1-3)[Man(a1-6)]Man(a1-6)[Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Man(a1-6)Man	Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc	Man(b1-4)GlcNAc	Man(b1-4)Man(b1-4)Man(b1-4)Man	Man(b1-4)Man(b1-4)Man(b1-4)Man(b1-4)Man	Man(b1-4)Man(b1-4)Man(b1-4)Man(b1-4)Man(b1-4)Man	Man6P(a1-3)Man(a1-3)Man(a1-3)Man(a1-2)Man	MurNAc(b1-4)GlcNAc	Neu(a2-3)Gal(b1-4)Glc	Neu(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu(a2-3)Gal(b1-4)GlcNAc6S(b1-3)Gal(b1-4)Glc	Neu(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc6S(b1-3)Gal(b1-4)Glc	Neu(a2-3)Gal6S(b1-4)[Fuc(a1-3)]GlcN(b1-3)Gal(b1-4)Glc	Neu(a2-6)Gal(b1-4)Glc	Neu4Ac5Ac(a2-3)Gal(b1-4)Glc	Neu4Me5Ac(a2-3)Gal(b1-4)Glc	Neu5Ac(a2-2)Glc	Neu5Ac(a2-3)4dGal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)6dGal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal	Neu5Ac(a2-3)Gal(b1-3)6dGalNAc	Neu5Ac(a2-3)Gal(b1-3)GalNAc	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)[Neu5Ac(a2-3)]Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)[Neu5Ac(a2-8)Neu5Ac(a2-3)]Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GalNAc6S	Neu5Ac(a2-3)Gal(b1-3)GalNAc6S(b1-4)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)]Man(a1-6)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-3)Gal(b1-3)GlcNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-3)GalNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)GalNAc	Neu5Ac(a2-3)Gal(b1-3)GlcNAc6S	Neu5Ac(a2-3)Gal(b1-3)[Fuc(a1-3)]GlcN(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc	Neu5Ac(a2-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc	Neu5Ac(a2-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Ac(a2-3)Gal(b1-3)[GlcNAc(b1-6)]GalNAc	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-3)Gal(b1-4)]GlcNAc	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GalNAc	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(b1-4)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-3)]Neu5Ac(a2-3)Gal(b1-4)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)]Man(a1-6)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)GalNAc(a1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)[Neu5Ac(a2-3)Gal(b1-3)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)GlcNAc6S	Neu5Ac(a2-3)Gal(b1-4)GlcNAc6S(b1-3)Gal6S(b1-3)GlcNAc6S	Neu5Ac(a2-3)Gal(b1-4)[3dFuc(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal(b1-4)[4dFuc(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal(b1-4)[6dTal(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]Glc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)GalNAc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)GalNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)[Neu5Ac(a2-3)Gal(b1-3)]GalNAc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc6S	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc6S(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc6S(b1-3)Gal6S(b1-3)GlcNAc6S	Neu5Ac(a2-3)Gal(b1-4)[Fuc2Me(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal(b1-4)[Qui(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal(b1-4)[Rha(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac(a2-3)Gal3S	Neu5Ac(a2-3)Gal6S(b1-4)GlcNAc	Neu5Ac(a2-3)Gal6S(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Ac(a2-3)Gal6S(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)Gal6S(b1-4)[Fuc(a1-3)]GlcNAc6S(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-3)GalNAc	Neu5Ac(a2-3)GalNAc(b1-4)GlcNAc	Neu5Ac(a2-3)GalNBz(b1-4)GlcNAc	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-3)Gal	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-3)GalNAc(b1-4)[Neu5Ac(a2-3)]Gal(b1-4)Glc	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-3)Glc	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-3)GalNAc	Neu5Ac(a2-3)[GalNAc(b1-6)]Gal(b1-4)Glc	Neu5Ac(a2-3)[Neu5Ac(a2-6)]GalNAc	Neu5Ac(a2-6)Gal	Neu5Ac(a2-6)Gal(b1-3)GalNAc(b1-4)Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-3)GlcNAc(b1-3)[Fuc(a1-3)[Gal(b1-4)]GlcNAc(b1-6)]Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-3)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(b1-4)Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Gc(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[GlcNAc(b1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)[Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-4)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]Glc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)GlcNAc(b1-3)]Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc	Neu5Ac(a2-6)Gal(b1-4)GlcNAc6S	Neu5Ac(a2-6)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Ac(a2-6)Gal(b1-6)GalNAc(b1-4)Gal(b1-4)Glc	Neu5Ac(a2-6)Gal6S	Neu5Ac(a2-6)Gal6S(b1-4)Glc	Neu5Ac(a2-6)Gal6S(b1-4)Glc4S	Neu5Ac(a2-6)GalNAc	Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc	Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc6S	Neu5Ac(a2-6)Glc	Neu5Ac(a2-6)Glc6S	Neu5Ac(a2-6)GlcNAc	Neu5Ac(a2-6)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)GlcNAc(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Ac(a2-6)GlcNAc6S	Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)[Neu5Ac(a2-3)]Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)[Neu5Ac(a2-8)Neu5Ac(a2-3)]Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-8)Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-6)Gal	Neu5Ac(a2-8)Neu5Ac(a2-6)Glc	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-6)Gal	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac	Neu5Ac(a2-8)Neu5Gc(a2-3)Gal(b1-4)GlcNAc	Neu5Ac(a2-9)Neu5Ac(a2-3)Gal(b1-4)Glc	Neu5Ac(b1-6)GalNAc	Neu5Ac(b2-3)Gal(b1-4)Glc	Neu5Ac(b2-6)Gal(b1-4)Glc	Neu5Ac(b2-6)Gal(b1-4)GlcNAc	Neu5Ac(b2-6)GalNAc	Neu5Ac9Ac(a2-3)Gal(b1-3)GlcNAc	Neu5Ac9Ac(a2-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc	Neu5Ac9Ac(a2-3)Gal(b1-4)GlcNAc	Neu5Ac9Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Ac9Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal	Neu5Ac9Ac(a2-6)Gal(b1-4)GlcNAc	Neu5Ac9Me(a2-3)Gal(b1-4)Glc	Neu5Gc(a2-3)Gal(b1-3)GlcNAc	Neu5Gc(a2-3)Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Gc(a2-3)Gal(b1-3)[Fuc(a1-4)]GlcNAc	Neu5Gc(a2-3)Gal(b1-4)Glc	Neu5Gc(a2-3)Gal(b1-4)GlcNAc	Neu5Gc(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Gc(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Gc(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc	Neu5Gc(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal	Neu5Gc(a2-3)Gal(b1-4)[Fuc(a1-4)]GlcNAc	Neu5Gc(a2-3)[GalNAc(b1-4)]Gal	Neu5Gc(a2-6)Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc	Neu5Gc(a2-6)Gal(b1-4)GlcNAc	Neu5Gc(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Gc(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Neu5Gc(a2-6)GalNAc	Neu5Gc(a2-8)Neu5Ac(a2-3)Gal(b1-4)GlcNAc	Neu5Gc(a2-8)Neu5Gc(a2-3)Gal(b1-4)GlcNAc	Neu5Gc(a2-8)Neu5Gc(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc	Neu5Gc(a2-8)Neu5Gc(a2-6)Gal(b1-4)GlcNAc	Neu5Gc(b2-6)Gal(b1-4)GlcNAc	Rha(a1-6)Glc	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc	Xyl(b1-2)[Man(a1-3)][Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Xyl(b1-2)[Man(a1-3)][Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	Xyl(b1-4)Xyl(b1-4)Xyl(b1-4)Xyl(b1-4)Xyl	Xyl(b1-4)Xyl(b1-4)Xyl(b1-4)Xyl(b1-4)Xyl(b1-4)Xyl	target	protein
0	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.293462	-1.316793	-0.860744	-1.211838	-0.335253	-1.127330	nan	-0.070201	-0.262080	-0.555655	-1.313299	-0.134637	0.317005	0.821230	nan	nan	-1.076498	nan	-0.068528	1.830645	-0.670428	0.691076	nan	0.276549	nan	nan	nan	nan	-1.170653	0.295167	-1.022834	0.401201	-1.117411	1.983681	-0.524707	-0.792918	0.556915	nan	0.281728	0.858230	0.750740	-0.326871	-0.490199	-0.019968	0.206706	-0.321137	-0.315444	nan	nan	-0.061100	nan	0.778259	nan	-1.122821	nan	-0.866634	0.898772	0.753109	nan	-0.570608	-0.370622	-0.214434	-0.083591	-0.202393	-0.234208	nan	nan	0.228359	-0.474023	-0.130498	-0.599990	-0.336374	1.158122	nan	-1.315247	-0.942153	1.501823	-0.334507	-0.251028	1.006794	-0.859050	nan	-1.982244	-0.676445	0.471350	0.036644	0.468515	-1.810952	nan	2.315709	0.199587	-0.505418	0.728243	0.026061	0.321230	-1.398725	nan	0.334855	-1.018204	nan	-0.620614	0.033777	1.862801	-0.315860	0.550124	nan	nan	-0.013552	-0.509655	-0.417772	-0.156735	-0.549880	-0.161960	-0.731572	-0.855801	-0.491026	nan	-0.555533	nan	nan	nan	nan	0.194983	0.512153	-0.996060	0.948385	0.618666	nan	0.454668	nan	nan	-0.321611	-1.673173	-0.605793	nan	nan	nan	nan	-0.062013	0.549441	nan	0.659781	-0.287028	nan	-0.555770	nan	nan	0.519453	nan	nan	nan	3.189814	-0.396293	nan	nan	1.141278	nan	-0.301479	-0.206983	nan	-0.661388	nan	-1.726143	-0.809143	0.359409	-1.036932	0.369392	0.387420	nan	-0.806718	-0.344472	-1.688612	-0.266892	0.345571	-1.348697	nan	0.730832	-1.557567	-0.635470	nan	-0.656538	-0.355969	nan	nan	-0.551805	0.196506	0.170764	-1.291785	nan	2.593879	1.790022	0.734081	-0.511112	2.920829	nan	-0.084440	-0.472743	-0.542256	-1.256444	-0.155544	nan	nan	0.177562	nan	0.291092	nan	-0.678792	-0.809929	0.826958	-0.520985	-1.125411	-0.545094	1.177174	0.473622	-0.096637	1.221971	-1.017643	0.182799	-0.348565	nan	0.205954	-0.934454	-0.408113	nan	-0.224584	0.662677	nan	nan	nan	nan	nan	0.002986	-0.262705	-0.366825	3.147103	-0.073753	0.506367	nan	1.229815	-0.763115	nan	nan	nan	0.013598	nan	nan	nan	-0.100337	2.391141	nan	-0.969170	nan	nan	nan	nan	-0.310658	nan	-0.444067	nan	-1.420922	-1.418361	1.068793	-0.134195	-0.333582	0.513704	0.443801	nan	0.361051	0.990320	0.345317	0.685388	-0.637602	-0.903252	nan	0.960072	-0.531655	0.205642	1.102914	0.194423	nan	-0.255641	nan	nan	3.527070	-1.013221	nan	-0.088100	-0.822409	nan	-0.699069	0.291485	-1.058475	-0.429073	nan	-0.484419	-0.822507	-0.778704	0.031066	0.102814	-0.013922	-0.104233	0.235792	nan	-0.692381	-0.371819	0.791884	-0.238029	-1.341460	0.567337	0.311051	-0.739965	1.503106	-0.777380	0.224688	0.204058	0.880339	1.169474	nan	-0.175544	-0.186680	-1.767021	nan	nan	0.074453	nan	nan	nan	nan	nan	0.861154	-0.275722	nan	0.760378	-0.122172	nan	0.866123	1.528204	-0.059112	nan	-1.266091	nan	1.919748	nan	-1.299860	nan	1.820241	0.909694	-0.394599	nan	nan	nan	0.417449	-0.030684	0.651839	-0.189469	2.275547	1.879986	0.034795	1.943014	0.132822	-1.320108	0.017783	nan	-0.536308	2.232935	0.993916	-0.398402	nan	0.613325	nan	0.657140	nan	0.083962	0.070875	-0.639347	0.963136	0.340265	-0.435801	1.314698	-0.707144	0.010442	nan	1.409104	1.312975	-0.647214	-0.378275	0.091274	nan	-1.098850	nan	-0.087915	-1.070024	-0.530146	1.161683	-0.544354	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.583946	nan	-0.205191	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.264347	-2.010649	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-1.293906	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-1.845928	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.252531	nan	nan	nan	nan	-0.370796	nan	nan	nan	nan	nan	-0.729693	nan	nan	-1.828778	1.019621	nan	1.163787	nan	-0.118932	nan	-0.209555	0.133655	-0.136596	0.335849	-0.673942	0.090147	-1.037059	nan	0.736757	nan	nan	nan	2.788943	0.725185	nan	0.424761	-0.145538	nan	0.841155	nan	nan	nan	-0.902281	-1.070313	-0.549892	0.370871	-1.507851	nan	-0.447876	0.387773	-0.060701	0.173437	-1.081284	-0.603117	-0.266577	-0.050777	0.402848	-0.523505	0.029309	-0.717797	-0.522505	0.114510	-0.156128	-0.597342	1.445367	-0.014696	-0.450986	0.580938	-0.866310	-0.152070	0.586291	-0.887315	-0.090753	-0.740103	0.386357	0.432027	nan	0.229075	nan	-0.104972	nan	nan	nan	0.158284	-0.600516	-1.672901	nan	1.439656	-0.051135	nan	nan	0.968090	-0.938656	-0.417570	nan	0.390362	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-1.561521	nan	0.509708	nan	0.121435	0.399651	-0.466956	nan	nan	nan	nan	nan	nan	-0.602359	nan	-0.592597	nan	nan	nan	0.189469	-1.230641	nan	0.034991	0.042089	-0.832484	nan	nan	0.599822	1.375992	-1.149122	nan	nan	nan	3.240981	nan	-1.280536	-0.512395	-0.006985	-0.368857	0.616198	-1.365917	nan	0.015448	nan	nan	nan	nan	-0.836073	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.938211	nan	-0.869553	1.554984	nan	nan	0.854212	0.392172	0.743144	nan	0.415767	-0.188451	-0.322970	0.679053	nan	-0.898535	nan	-0.568447	0.525800	nan	-0.389333	-0.944222	-0.021667	nan	-1.095452	1.406847	nan	1.647619	-0.435847	nan	0.082123	nan	nan	-1.215352	0.718515	1.217300	-0.435636	-0.663364	nan	-0.429130	0.201278	-0.509222	-0.106666	nan	-1.167387	0.340328	nan	0.740803	nan	-0.364258	-1.250190	-0.878616	nan	0.077389	2.517735	nan	nan	-0.019361	1.491176	-0.303617	-0.672405	0.317837	0.332005	0.333930	-2.222108	0.120233	nan	-0.585597	-1.454731	0.470726	nan	nan	nan	nan	nan	-0.955375	-1.255837	0.372987	nan	1.995031	nan	-0.188035	-0.179925	nan	nan	-0.340091	-0.871518	1.115365	nan	nan	nan	nan	nan	nan	0.867050	-1.056383	nan	nan	0.412622	0.198457	nan	nan	nan	nan	-1.028291	-0.761279	0.779121	nan	-0.633336	-0.127695	nan	nan	0.175828	nan	0.220810	1.906694	-0.769774	1.141544	0.148053	-0.493245	nan	-0.335814	-1.279935	0.793184	-0.408281	0.572545	nan	-1.045371	0.655827	-0.839004	-0.593631	nan	0.212700	-1.616937	-0.278947	0.252150	nan	-0.421107	-1.389338	-1.094746	-0.881483	3.894794	1.396125	1.640365	nan	-1.230901	1.346605	nan	0.082465	-0.710069	-1.089082	-1.185716	1.357356	nan	nan	nan	nan	nan	0.179706	0.778520	0.790346	nan	nan	nan	0.492500	-0.722607	nan	0.607282	nan	-1.436072	0.068759	-1.488441	1.208220	-0.586152	-0.463199	nan	nan	0.490009	-1.627624	-0.364351	-0.514077	nan	nan	-0.204468	nan	nan	nan	nan	nan	nan	nan	0.658498	nan	nan	nan	nan	-0.091533	-1.166208	0.626637	nan	-1.109417	nan	nan	-0.388218	nan	0.455506	nan	0.443298	-0.000956	-1.520597	-0.171781	0.156926	nan	nan	nan	nan	0.419963	nan	-0.225532	0.185515	-1.111053	0.759797	-0.801270	-0.285606	nan	nan	nan	nan	nan	nan	AADSIPSISPTGIITPTPTQSGMVSNCNKFYDVHSNDGCSAIASSQHVDLSSLYKWNPAIKTDCSGLQASVYVCIGILTTSMSTTSKLPTTTSKPSTTSKPPTGITTPTPTQSGMVKNCNKFYDVHSGDGCSAIASSQKVNLSSFYLWNPAVKTDCSGLQASVFVCIGTLTTSMSTTTSKLPTTTSKPSTTSKPPTGIITPTPTQNGMVKNCNKFYDVHSGDGCSAIASSQKVNLSSFYLWNPAVKTDCSGLQASVFVCVGTATTTTAAGITTPTPTQSGMVSGCNKFYDVHAGDGCSAIASSQKIALSSLYKWNPAVKTDCSGLQASVYICIGVGNAAAARVTGALEQKLISEEDLNSAVDHHHHHH	TAL6-4LysM
1	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.210341	-0.317977	-0.535696	-0.428562	0.304758	-0.599696	nan	-0.514044	-0.621800	0.110234	1.704315	-0.342968	-0.241699	-0.094321	nan	nan	0.548443	nan	-0.164111	-0.301651	-0.286659	5.996096	nan	0.195205	nan	nan	nan	nan	-0.572861	-0.078468	-0.597313	-0.271587	-0.185770	-0.477049	-0.148226	0.273879	3.048219	nan	1.536058	-0.388298	0.720448	-0.154050	0.145095	-0.120509	-0.642615	-0.358526	-0.401998	nan	nan	-0.453027	nan	0.746013	nan	0.881687	nan	-0.673569	-0.409280	-0.241929	nan	-0.084825	-0.452808	-0.526735	0.338334	-0.350185	-0.280775	nan	nan	-0.219135	-0.214087	-0.278396	-0.039948	0.319400	0.767513	nan	0.321303	0.054521	-0.004854	-0.394323	-0.397670	-0.363056	0.408759	nan	-0.538161	-0.072649	-0.305646	-0.598274	-0.146399	-0.588251	nan	0.127787	-0.389585	-0.544289	-0.273656	0.496878	-0.562748	0.696372	nan	0.292971	-0.439380	nan	-0.487665	-0.387022	-0.723142	-0.716450	1.611207	nan	nan	-0.422152	0.319929	-0.606341	-0.160275	-0.494404	0.049719	1.054716	-1.226240	-0.689861	nan	-0.394123	nan	nan	nan	nan	0.209877	-0.632032	-0.279757	0.120900	-0.035907	nan	-0.393659	nan	nan	-0.163429	-0.735791	-0.050865	nan	nan	nan	nan	0.153107	0.159939	nan	0.114519	-0.238581	nan	0.243765	nan	nan	-0.205723	nan	nan	nan	-0.169214	-0.537188	nan	nan	0.108785	nan	-0.017564	-0.278367	nan	-0.706053	nan	-0.421375	-0.637758	0.325945	-0.185604	-0.206844	-1.225292	nan	-0.201082	-0.284139	-0.316140	1.195916	-0.161080	-0.551468	nan	-0.120453	3.751031	-0.044979	nan	-0.627522	0.714630	nan	nan	-0.307018	-0.428596	3.366465	-0.475412	nan	-0.156273	-0.273549	-0.257248	-0.077096	-0.238332	nan	-0.229461	0.020193	-0.151334	-0.297753	-0.169976	nan	nan	-0.071467	nan	-0.118106	nan	-0.315901	-0.184823	0.176021	-0.056910	-0.571056	0.493649	0.506835	1.096384	-0.285038	-0.328707	0.059036	1.714136	-0.023349	nan	1.438082	0.560742	-0.064961	nan	0.293321	-0.738018	nan	nan	nan	nan	nan	0.438784	-0.647943	-0.911864	-0.147095	-0.076763	-0.105693	nan	-0.291598	-0.332561	nan	nan	nan	-0.471061	nan	nan	nan	-0.623435	-0.423218	nan	0.335238	nan	nan	nan	nan	-0.458750	nan	-0.458470	nan	-0.620907	-0.433047	-0.746319	0.103050	-0.424069	-0.219640	3.941640	nan	-0.204654	0.140085	0.004276	0.558532	0.196721	-0.263195	nan	-0.932223	-0.267338	-0.189840	-0.193638	-0.518012	nan	-0.587022	nan	nan	0.464959	2.988462	nan	0.522682	1.023056	nan	2.066318	3.411378	-0.375306	0.986010	nan	-0.209109	-0.503419	0.011710	-0.336139	5.985190	1.319522	1.268927	2.710962	nan	0.528672	4.452175	1.298506	4.112301	2.141467	1.655698	4.640982	0.770460	0.433840	3.938218	2.332191	1.565793	2.655383	1.949440	nan	-0.135356	-0.320277	0.660683	nan	nan	-0.071170	nan	nan	nan	nan	nan	0.091177	-0.504007	nan	-0.307391	-0.085958	nan	0.039727	-1.266585	-0.148673	nan	-0.367992	nan	-0.308856	nan	-0.266432	nan	-0.303892	-0.416294	-0.132050	nan	nan	nan	-0.506776	-0.075666	-0.185265	-0.316894	-0.323859	-0.596264	-0.376123	-0.281852	-0.326506	-0.777421	-0.576470	nan	-0.662114	-0.504215	-0.713073	-0.449993	nan	-0.243328	nan	-0.628874	nan	-0.574322	-0.006513	-0.048031	0.014492	-0.096467	0.329244	-0.444026	-0.496267	0.571160	nan	-0.019903	0.037512	3.418474	-0.062730	-0.072516	nan	-0.267884	nan	-0.340286	-0.693033	-0.664153	-0.578903	-0.260458	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.029600	nan	0.085726	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.061387	-0.230035	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.068152	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.146414	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.929766	nan	nan	nan	nan	0.021350	nan	nan	nan	nan	nan	1.188295	nan	nan	-0.613549	-0.792706	nan	-0.305427	nan	-0.191716	nan	-0.786313	-0.728967	-1.106288	-0.380770	-0.522477	0.091364	0.294415	nan	-0.603658	nan	nan	nan	-0.374202	1.311709	nan	-0.318125	-0.164150	nan	0.031533	nan	nan	nan	-0.276207	-0.196006	-0.642186	-0.226948	1.353726	nan	-0.425933	-0.004902	-0.618264	-0.116827	-0.626297	-0.454391	0.076760	-0.352582	-0.546664	-0.255336	3.022814	0.405144	-0.859338	-0.214939	0.698437	-1.061306	-0.268759	0.053256	-0.432154	-0.453639	-0.132935	0.249749	-0.117152	-0.124793	-0.667753	-0.560969	0.734671	-0.363380	nan	-0.723359	nan	-0.541242	nan	nan	nan	0.711890	-0.274278	-0.208130	nan	-0.146078	-0.305149	nan	nan	-0.002285	-0.312072	-0.194684	nan	0.135019	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.192462	nan	-0.880676	nan	0.222977	-0.261812	0.026415	nan	nan	nan	nan	nan	nan	-0.099068	nan	-0.432871	nan	nan	nan	-0.006577	-0.482137	nan	-0.986270	-0.570354	0.842646	nan	nan	-0.650632	-0.958361	0.133986	nan	nan	nan	-0.022822	nan	1.718850	-0.085626	0.139780	-0.211927	-0.498004	-0.763792	nan	-0.160861	nan	nan	nan	nan	-0.247478	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.334203	nan	-0.476253	-0.224135	nan	nan	-0.333129	0.303401	-0.283198	nan	-0.334101	-0.094590	-0.467862	5.370085	0.155618	0.092594	nan	-0.252477	-0.121639	nan	0.011680	0.200834	0.753740	nan	-0.549464	-0.194579	nan	-0.238250	-0.348041	nan	-0.429747	nan	nan	0.044244	-0.474801	0.341388	-0.728712	0.283256	nan	-0.431491	-0.329709	0.096213	-0.712527	nan	0.042465	-0.115827	nan	nan	nan	-0.417758	-0.931021	-0.432294	nan	-0.022245	1.624166	nan	nan	-0.539692	4.099614	-0.397088	0.489274	-0.373275	-0.136173	-0.263554	-0.192120	-0.107791	nan	0.089840	-0.210930	-0.489136	nan	nan	nan	nan	nan	-0.535536	-1.350794	-0.441527	nan	0.271697	nan	0.143828	0.502451	nan	nan	-0.389743	0.160502	-0.443902	nan	nan	nan	nan	nan	nan	-0.759362	-0.672274	nan	nan	-0.763808	0.043080	nan	nan	nan	nan	-0.324763	-0.735311	0.606204	nan	-0.142990	-0.537923	nan	nan	0.024373	nan	-0.232817	-0.253269	-0.167730	-0.302303	-0.272344	-0.465127	nan	-0.331235	-0.337413	-0.337699	-0.048925	-0.204174	nan	-0.112222	-0.372519	-0.381540	-0.629240	nan	-0.212922	-0.068444	-0.275976	0.274100	nan	-0.269315	-0.020473	0.059676	3.188752	-0.179521	2.341252	-0.716474	nan	-0.502049	0.044380	nan	-0.190638	0.066739	-0.347539	-0.387395	-0.577035	nan	nan	nan	nan	nan	-0.379801	-0.632533	-0.457350	nan	nan	nan	-0.601558	-0.895900	nan	-0.141286	nan	-0.272919	0.387567	-0.526855	-0.847587	-0.384928	-0.715208	nan	nan	-0.552275	-0.435920	0.673410	-0.806533	nan	nan	-0.316786	nan	nan	nan	nan	nan	nan	nan	-0.765163	nan	nan	nan	nan	-0.665418	-0.547333	-0.188309	nan	2.013600	nan	nan	-0.285953	nan	-0.114993	nan	-0.318728	-0.609542	-0.444485	0.288337	-0.236235	nan	nan	nan	nan	-0.087587	nan	-0.230830	-0.591006	-0.542732	-0.645822	-0.630642	-0.083163	nan	nan	nan	nan	nan	nan	AAFFSLVVLLALLPFGIHASALPSTELTPRVNPNLPGPNDVFVGFRGTNNIVGAIYEKDGIKDTPLNNGADAEMGPGSYITDGLFTARIFASGSAGNNPGTTATVCKIYASGVQWRGAPKGYIPVAYFGNGPTAEQRRQTYLRTLGLPGTAIRFGNLNTNPGQGPTAPRQMVVPPGPLRSLLKASCFPAADTRLPQTLHYFEKRVDWNIITAADSQTWGH	rCnSL-proA
2	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.141335	-0.048689	0.028283	-1.017600	-0.871024	-1.540761	nan	-0.350449	-0.652191	0.716365	0.612228	-1.258691	0.237488	0.422113	nan	nan	-1.543964	nan	-1.187212	0.457266	0.480527	0.420569	nan	0.614918	nan	nan	nan	nan	-0.839644	-0.770383	4.683932	-0.064917	1.149108	-0.565169	1.387117	1.495458	1.109946	nan	0.655759	-1.040241	0.724848	0.654293	-0.034218	0.338654	-0.844789	1.889215	-0.715226	nan	nan	0.076582	nan	2.555725	nan	0.208505	nan	0.169653	1.500775	0.835543	nan	-0.889909	-1.078300	0.470129	-0.124160	-0.552058	0.254316	nan	nan	-0.227861	0.123434	-0.957585	-0.863318	-0.497218	-0.592302	nan	1.597832	0.337272	-0.571593	-1.002809	-0.527701	0.173993	3.104163	nan	-0.213374	nan	-0.016564	0.825037	-0.161087	-0.145769	nan	-0.200245	-1.128504	-0.354246	-0.495097	0.349565	1.033642	-0.775093	nan	0.738992	-0.331531	nan	-0.729900	-0.846219	-0.980832	1.013321	0.155313	nan	nan	-0.715678	0.176707	0.387436	0.200393	-0.876682	0.390241	1.570174	0.887831	0.107227	nan	0.171433	nan	nan	nan	nan	-0.335483	0.825236	-0.374665	0.125660	0.760958	nan	-0.626154	nan	nan	0.906766	-0.372149	2.642953	nan	nan	nan	nan	0.145219	-1.157278	nan	0.209840	0.537053	nan	0.993735	nan	nan	1.185427	nan	nan	nan	0.931615	-0.390090	nan	nan	-0.388097	nan	-0.590225	-0.595967	-0.495262	-0.196759	nan	-0.563083	-0.640716	3.466913	2.789688	0.992895	0.369458	nan	-0.763190	-0.982501	-0.101580	1.548296	-1.152511	-0.128297	nan	-1.036687	0.993182	1.360269	nan	0.240559	2.055367	nan	nan	0.900653	-0.269334	-0.010582	-0.067021	nan	-0.879200	-0.190946	-0.551266	0.084391	-0.051376	nan	-0.409344	1.010344	0.077779	-0.620924	0.437636	nan	nan	-0.798348	nan	-0.731754	nan	0.211020	-0.673579	0.451430	-0.152162	0.099131	0.484088	1.129303	0.815293	1.591959	-0.068063	0.484870	-0.071728	1.148531	nan	5.611608	-1.076119	-0.401547	nan	-0.768734	-0.215077	nan	nan	nan	nan	nan	-0.879382	-1.147433	-0.090986	0.093784	-0.532454	0.064582	nan	0.264495	-1.390338	nan	nan	nan	0.182229	nan	nan	nan	3.096448	0.648753	nan	-0.774947	nan	nan	nan	nan	-0.670909	nan	-1.083155	nan	0.658986	-0.098134	-1.495887	0.297815	1.203526	-0.744118	1.048740	nan	0.599308	-0.225759	-1.123019	1.807711	-0.213169	-1.220352	nan	-0.072945	-0.554981	1.673273	0.216192	-0.498000	nan	0.472995	nan	nan	-0.518564	0.027248	nan	-0.310421	0.598330	nan	-0.417603	-0.250864	-0.581462	-0.573974	nan	-0.492076	0.173055	-0.380729	-0.087446	0.700929	2.037591	1.555403	1.535190	nan	-0.395869	0.926895	1.394170	1.808713	2.517612	3.269472	2.270964	2.197590	1.300799	1.714303	3.905954	1.995075	0.339238	-0.398401	nan	-0.717286	-1.151917	-0.738147	nan	nan	0.600467	nan	nan	nan	nan	nan	0.525111	-0.192369	nan	-0.203349	0.419092	nan	0.429460	-0.923743	-0.487743	nan	-1.051212	nan	-0.670214	nan	0.514861	nan	0.494058	-0.822357	0.443901	nan	nan	nan	3.679210	-0.047971	0.489464	-0.164600	0.603060	0.435522	-0.506143	-1.142534	-1.278513	-0.028601	0.363345	nan	-0.109694	1.009996	0.781620	0.522848	nan	0.000915	nan	0.326645	-0.515516	-0.223894	nan	-0.411156	0.730003	-0.843211	-1.058670	1.370434	-0.475073	-0.825378	nan	0.185191	-0.796588	-0.867760	-0.774234	-1.301059	nan	-0.454536	nan	-1.462383	-0.649293	-0.559610	-0.462466	0.953470	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.184253	nan	-1.188183	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.587275	0.176616	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.226841	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.364115	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-1.100772	nan	nan	nan	nan	-0.575073	nan	nan	nan	nan	nan	0.668801	nan	nan	-1.077393	-0.668994	nan	-0.723625	nan	-0.217599	nan	-0.792700	-0.916332	0.081845	1.468745	-0.892370	0.069646	0.177320	nan	-1.058256	nan	nan	nan	-0.622087	-0.727465	nan	-1.108628	1.460117	nan	-0.122677	nan	nan	nan	-1.349787	-0.894629	-0.588259	0.082347	0.953946	nan	-0.813898	-1.017165	-0.153106	-0.295210	-1.245724	0.116423	-0.320367	0.424547	0.978872	-0.330253	2.186425	0.477924	-0.456296	-0.017359	0.510488	1.956776	1.126133	3.226699	0.602039	-1.005354	-0.624235	-0.581631	-0.907495	0.271080	-0.012440	-0.709801	-1.029425	-0.953545	nan	-1.174501	nan	-0.557060	nan	nan	nan	-0.561542	0.034974	0.168207	nan	0.051790	-0.579770	nan	nan	-0.642614	-0.001472	-0.654685	nan	0.769802	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.902390	nan	0.431743	nan	0.137663	-0.618213	1.133564	nan	nan	nan	nan	nan	nan	-1.054152	nan	-1.160963	nan	nan	nan	1.221059	0.520033	nan	-0.805138	-1.512611	-0.708850	nan	nan	-0.990863	-1.480671	0.974897	nan	nan	nan	0.130424	nan	0.367533	-0.420475	-0.421585	-0.410854	-0.692633	-0.277207	nan	-0.516160	nan	nan	nan	nan	0.740449	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.346629	nan	-0.763731	0.491866	nan	nan	-0.369327	0.854738	-0.385137	nan	-0.142725	-0.920891	-0.192656	0.302544	nan	-0.515233	nan	-1.160781	-0.806743	nan	-0.890371	0.015838	-1.254571	nan	-0.416332	-0.256326	nan	-0.386644	0.442694	nan	-1.056443	nan	nan	-0.099207	0.015878	-0.467975	0.012557	0.920586	nan	-0.724094	-0.300478	-1.077420	0.080169	nan	0.009904	-0.940578	nan	0.406078	nan	-1.348789	-1.178166	0.366113	nan	-0.615617	-0.057792	nan	nan	-0.964989	1.396847	1.421386	0.323981	-0.290309	-0.593175	-1.272687	0.764037	-1.013082	nan	-0.572224	0.144831	-0.627496	nan	nan	nan	nan	nan	-0.523728	2.242855	-0.529255	nan	-0.000575	nan	-0.070429	1.568826	nan	nan	2.321792	0.167485	-1.036465	nan	nan	nan	nan	nan	nan	-0.680242	-0.675306	nan	nan	-1.048313	-0.789362	nan	nan	nan	nan	-0.635699	-0.450291	0.843699	nan	0.222346	-1.554352	nan	nan	0.439390	nan	-0.114403	-0.451667	-0.521770	-0.244926	0.156868	-0.928180	nan	-0.618409	0.483767	0.381458	-0.329480	1.668495	nan	-0.280346	-1.050066	-0.891082	-1.133454	nan	-0.285970	1.051619	0.992757	-1.100358	nan	-0.213941	-0.823419	-1.168836	1.737089	-0.349192	0.793185	0.305781	nan	0.246530	0.206354	nan	-1.080515	0.255455	-0.279725	-1.322618	1.060777	nan	nan	nan	nan	nan	-0.370770	-1.167221	-0.769847	nan	nan	nan	-0.597977	-0.712772	nan	-0.380666	nan	0.241274	-0.035084	-0.254492	-0.756890	-0.073090	0.406664	nan	nan	-0.735348	-0.756698	0.119479	0.774519	nan	nan	0.749350	nan	nan	nan	nan	nan	nan	nan	-0.976446	nan	nan	nan	nan	0.331058	-1.136037	-1.544240	nan	-1.291898	nan	nan	-0.505057	nan	-0.716022	nan	-0.884804	-0.071245	-0.464590	1.883028	0.707221	nan	nan	nan	nan	0.797868	nan	0.493683	1.662689	-0.703489	-1.168415	-1.297091	1.005849	nan	nan	nan	nan	nan	nan	AANEADYQAKLTAYQTELARVQKANADAKAAYEAAVAANNAANAALTAENTAIKKRNADAKADYEAKLAKYQADLAKYQKDLADYPVKLKAYEDEQASIKAALAELEKHKNEDGNLTEPSAQNLVYDLEPNANLSLTTDGKFLKASAVDDAFSKSTSKAKYDQKILQLDDLDITNLEQSNDVASSMELYGNFGDKAGWSTTVSNNSQVKWGSVLLERGQSATATYTNLQNSYYNGKKISKIVYKYTVDPKSKFQGQKVWLGIFTDPTLGVFASAYTGQVEKNTSIFIKNEFTFYDEDGKPINFDNALLSVASLNREHNSIEMAKDYSGKFVKISGSSIGEKNGMIYATDTLNFKQGEGGSRWTMYKNSQAGSGWDSSDAPNSWYGAGAIKMSGPNNYVTVGATSATNVMPVSDMPVVPGKDNTDGKKPNIWYSLNGKIRAVNVPKVTKEKPTPPVKPTAPTKPTYETEKPLKPAPVAPNYEKEPTPPTRLEHHHHHH	AntigenI/IIA3VP1
3	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.951637	-0.590611	-0.209295	-1.768618	0.072966	-0.120822	nan	-0.588345	-0.073485	-1.229778	-0.903323	-0.233609	-0.595171	0.213082	nan	nan	0.974062	nan	-1.468367	0.905727	-1.174894	-2.406395	nan	0.448905	nan	nan	nan	nan	-0.674998	-0.124440	0.066959	0.310361	0.072212	1.268628	-0.376546	0.429747	-1.153319	nan	0.463288	-0.313873	-1.396695	0.083592	0.064160	-0.205204	0.428165	3.118173	0.291483	nan	nan	-0.195754	nan	0.008242	nan	-0.207449	nan	-0.762228	2.678329	1.621970	nan	1.821577	0.063804	-0.395757	1.447813	-0.384916	0.326316	nan	nan	0.612385	-0.805186	0.209411	1.378479	-0.506181	-1.232264	nan	-0.132982	0.722559	0.490832	1.196810	-0.907167	0.478301	1.704462	nan	-0.999434	-0.723832	-0.509363	-0.694436	-0.889037	0.955400	nan	0.687000	-0.756971	-0.872953	-0.404418	0.346657	-0.391230	-1.064233	nan	0.203522	1.125154	nan	-0.723746	0.557047	0.179450	0.420203	-2.017785	nan	nan	-0.762109	-0.144362	-0.337741	1.582901	1.535322	0.713354	-0.108216	-0.239374	-0.198454	nan	0.348999	nan	nan	nan	nan	0.625678	-0.409209	0.367381	-0.825724	0.672665	nan	0.406999	nan	nan	-0.013477	-0.382688	0.693050	nan	nan	nan	nan	-0.638029	0.048011	nan	0.506819	-0.760823	nan	0.225662	nan	nan	-0.382483	nan	nan	nan	2.991649	0.001304	nan	nan	-0.873890	nan	-0.452589	-0.943522	nan	-0.439595	nan	-0.376261	0.127424	-0.832081	0.586475	-1.257440	1.501641	nan	-0.260007	2.223311	0.538368	0.080427	-1.018306	-0.541649	nan	-0.417671	-0.687471	0.186877	nan	0.464817	-0.972353	nan	nan	2.693062	0.404590	-0.586090	-0.008127	nan	-1.603526	0.524593	-0.670450	-0.191908	0.459009	nan	-0.343084	-2.145714	-0.299270	-1.582209	-0.336853	nan	nan	-0.503355	nan	-0.686884	nan	-0.122241	0.950281	-1.168348	3.656594	0.699332	-0.231499	-0.060755	-0.157388	1.473764	-0.601700	-0.570523	0.355733	-0.913045	nan	0.008199	-1.311791	-0.428372	nan	-1.391807	-0.728623	nan	nan	nan	nan	nan	0.847592	-0.120010	-1.012224	-0.288547	0.801761	0.148875	nan	-0.760349	0.447414	nan	nan	nan	-1.195392	nan	nan	nan	1.020333	-0.669842	nan	-0.094799	nan	nan	nan	nan	-0.503495	nan	0.532867	nan	-0.396656	0.915846	-1.312695	0.667610	-1.078118	-0.744122	-0.177299	nan	0.066577	-0.341433	0.478667	-1.432792	-1.239424	0.551174	nan	-0.218031	0.569050	-1.586408	-0.133160	-0.648386	nan	0.445212	nan	nan	0.340408	-0.570270	nan	-1.108098	1.237671	nan	-0.309745	-1.184158	-0.993529	0.536947	nan	-0.573462	-1.816202	-0.431790	-1.181353	-0.047435	0.711572	0.687726	-0.076688	nan	0.635663	-0.179234	-0.033232	2.093702	1.773395	-0.763503	0.045718	-1.114337	0.184397	0.687753	0.243150	1.432827	0.100575	0.642413	nan	0.867767	0.941375	-0.192011	nan	nan	1.147492	nan	nan	nan	nan	nan	-1.597631	-0.255916	nan	2.004675	-0.644327	nan	1.093860	0.256672	1.478630	nan	0.856145	nan	0.215771	nan	0.692617	nan	-0.867506	1.857473	0.433973	nan	nan	nan	2.958841	-0.685457	0.702930	-0.547387	2.508914	2.997883	0.092527	0.716433	-1.321049	-0.839434	-1.524387	nan	0.457076	0.947794	0.000163	-0.018150	nan	2.440934	nan	-1.350483	nan	0.453706	-0.793529	1.284174	-0.315127	0.375951	-0.541687	-0.839709	-0.240699	0.272568	nan	1.070937	-1.474703	1.817841	-1.491648	0.992186	nan	-0.302666	nan	-0.701536	0.679663	-0.003395	-1.076716	0.157181	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.578731	nan	-0.352899	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.776132	-0.389718	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.771739	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.128541	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.743590	nan	nan	nan	nan	1.114156	nan	nan	nan	nan	nan	0.562752	nan	nan	2.024226	0.145742	nan	-0.284101	nan	0.821275	nan	-0.903356	0.444561	-1.305724	-1.485496	-0.008224	-0.364553	0.905344	nan	-0.296444	nan	nan	nan	0.420031	0.127704	nan	-1.172369	-0.838201	nan	-0.558982	nan	nan	nan	1.148908	0.222792	0.450762	-1.160263	1.142438	nan	-0.275461	-0.653957	-0.361974	0.060989	0.267777	0.020739	-1.200785	0.822755	1.762667	0.291381	0.251733	0.195232	-0.539022	0.511997	-0.349578	-0.372837	-0.734474	1.317505	2.310868	-0.562195	0.441697	0.159291	-0.871995	-0.152614	-0.350767	-0.097614	-0.043341	1.903397	nan	-0.187887	nan	-1.077811	nan	nan	nan	-0.966938	-0.196707	0.249518	nan	0.737324	-0.323772	nan	nan	1.001472	-1.185256	1.316377	nan	0.253259	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.312635	nan	-1.176665	nan	-0.558890	0.209981	1.879744	nan	nan	nan	nan	nan	nan	2.709582	nan	-0.997819	nan	nan	nan	-1.131879	-0.128272	nan	-0.347904	-0.014177	-0.419786	nan	nan	-0.092632	0.122311	-0.099242	nan	nan	nan	-0.021406	nan	2.524406	-0.725861	0.135504	-0.273553	0.171989	-0.108189	nan	0.529707	nan	nan	nan	nan	0.385468	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.154296	nan	-1.434264	-0.741321	nan	nan	-0.799795	-0.659313	0.648770	nan	-0.539046	0.460526	0.806580	-1.382269	nan	1.138331	nan	-0.457176	2.365913	nan	0.273375	-0.412471	-0.031418	nan	1.222739	-0.755973	nan	1.023234	-1.626629	nan	0.886790	nan	nan	0.865350	-0.806674	-0.572708	-0.585570	0.220300	nan	0.193585	-0.417197	0.049115	-0.682798	nan	-0.980696	-0.720602	nan	0.484905	nan	0.012646	-1.164252	-0.146854	nan	-0.540276	-0.657763	nan	nan	1.756741	-0.398303	0.926273	-0.648273	-0.274664	0.025699	-0.542532	-1.250211	0.786932	nan	-1.452149	-1.140013	-0.459964	nan	nan	nan	nan	nan	-0.196312	-0.793960	0.168743	nan	-0.591144	nan	0.057754	-0.077485	nan	nan	0.957655	-0.367007	0.058728	nan	nan	nan	nan	nan	nan	0.427600	-1.041480	nan	nan	0.267018	-0.544599	nan	nan	nan	nan	-1.281566	-1.083673	1.671810	nan	0.398731	-0.907107	nan	nan	-0.730259	nan	0.029384	-0.076599	-0.567314	-0.141536	0.695219	0.741668	nan	0.064321	-0.593798	-0.013122	0.076119	-0.890878	nan	0.838301	0.106119	-0.010285	-1.406960	nan	0.287860	-0.828776	0.005115	-1.095572	nan	1.521757	1.366456	1.285267	1.615747	-0.316575	0.003150	-0.387150	nan	2.128922	-1.094625	nan	-0.108200	-0.092169	-1.033255	-0.007605	0.283393	nan	nan	nan	nan	nan	-0.324903	-0.721049	1.989544	nan	nan	nan	-0.899679	-1.003498	nan	-0.044499	nan	-0.252589	0.051698	-1.510983	-1.090636	-0.398503	2.166661	nan	nan	-0.138793	-0.818538	-0.053375	1.195357	nan	nan	0.372549	nan	nan	nan	nan	nan	nan	nan	-0.396403	nan	nan	nan	nan	0.125583	-0.369397	-1.092181	nan	-1.111521	nan	nan	-0.339356	nan	-0.841087	nan	-1.239822	-1.249646	-0.604720	-1.838380	0.247118	nan	nan	nan	nan	0.191007	nan	-0.669702	-0.626241	-1.722196	-0.429002	-0.454920	0.217270	nan	nan	nan	nan	nan	nan	AASKLGVPQPAQRDQVNCQLYAVQPNDNCIDISSKNNITYAQLLSWNPSLSSTCSNLASLNSSSICVSNPKGTFSISSNTVGATDIATTTAPVPSPTLDQTTSRCAKYYQVSDGDDCSHLTAQFAITLKDFIFLNSEVWQNCTNLELGYYYCVEPVGYISTYPGYLPTATTKPFNQTSATSLPYDGDPWARFSSNSSVIPIANGTRVDCYSYVYVKNLTENLFADCWNMASMYEITREELVLWNPSLGNDGSSGSNLSGAEASQVASSIAIPTTAPSSITTNLYTYPCTVAANISYCVALVSSTGALPTNTAPPGPHASGEISNCTAWFAPEAYDTCKSILDIFEMSFANFYKMNPSVGPDCSGLAVGTNYCVSTYPNGEDPNDDWDGDDSIPSISPTGIITPTPTQSGMVSNCNKFYDVHSNDGCSAIASSQHVDLSSLYKWNPAIKTDCSGLQASVYVCIGILTTSMSTTSKLPTTTSKPSTTSKPPTGITTPTPTQSGMVKNCNKFYDVHSGDGCSAIASSQKVNLSSFYLWNPAVKTDCSGLQASVFVCIGTLTTSMSTTTSKLPTTTSKPSTTSKPPTGIITPTPTQNGMVKNCNKFYDVHSGDGCSAIASSQKVNLSSFYLWNPAVKTDCSGLQASVFVCVGTATTTTAAGITTPTPTQSGMVSGCNKFYDVHAGDGCSAIASSQKIALSSLYKWNPAVKTDCSGLQASVYICIGVGNAAAARVTGAAASFLEQKLISEEDLNSAVDHHHHHH	TAL6-6LysM
4	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.352142	0.244700	-0.461016	-0.234837	-0.325556	-0.052849	nan	-0.291694	-0.201436	0.068008	0.284881	-0.207676	-0.509240	0.261744	nan	nan	-0.461065	nan	0.281879	0.314454	-0.547046	0.065653	nan	-0.206745	nan	nan	nan	nan	0.069357	-0.440782	-0.096734	-0.352217	-0.313786	-0.054417	0.220397	0.000593	-0.468453	nan	-0.642583	-0.213253	0.004967	-0.418988	0.019801	0.348693	-0.167535	-0.072314	-0.415624	nan	nan	-0.007247	nan	0.493524	nan	-0.049574	nan	0.356735	-0.231039	0.687253	nan	0.039245	0.289550	-0.017010	0.552749	0.253952	0.024845	nan	nan	-0.092765	-0.050599	-0.253661	-0.034002	-0.640639	-0.332807	nan	-0.456518	0.013717	-0.257121	-0.423334	0.238793	1.040417	-0.252777	nan	-0.459521	8.334515	-0.000602	0.304265	-0.082544	-0.571446	nan	0.359063	-0.372296	-0.115188	-0.467803	-0.495880	-0.284533	-0.135805	nan	0.521431	-0.180462	nan	-0.311932	-0.437642	-0.499129	-0.580729	-0.433832	nan	nan	-0.622161	0.178139	-0.593969	-0.479843	-0.404232	0.426017	0.186881	-0.169694	-0.266457	nan	-0.062682	nan	nan	nan	nan	-0.087707	-0.428183	0.186681	-0.260413	-0.097686	nan	-0.084458	nan	nan	-0.236920	0.000047	-0.328665	nan	nan	nan	nan	-0.050001	-0.388068	nan	0.096289	-0.713843	nan	0.718743	nan	nan	-0.192620	nan	nan	nan	0.140275	-0.483201	nan	nan	19.126138	nan	0.447124	-0.091157	nan	0.236213	nan	0.958505	-0.137814	0.299341	-0.456594	-0.101001	0.394345	nan	-0.381016	0.230053	-0.413905	0.708616	-0.731750	0.055039	nan	-0.425287	-0.560030	-0.214257	nan	1.285163	-0.247194	nan	nan	0.738753	0.260701	0.550376	-0.507198	nan	0.095765	0.431800	0.055439	0.251386	-0.593213	nan	-0.057296	-0.520796	-0.694564	-0.697927	-0.489050	nan	nan	-0.148911	nan	-0.500282	nan	0.060701	0.216923	-0.414411	2.763139	0.261430	-0.152436	-0.152840	-0.324842	0.024281	-0.436718	0.029816	0.086215	0.558052	nan	0.752209	0.016113	-0.574978	nan	0.183228	0.028883	nan	nan	nan	nan	nan	-0.558211	0.161656	-0.273586	-0.287044	-0.203313	0.303259	nan	0.610638	-0.080314	nan	nan	nan	-0.228164	nan	nan	nan	0.188941	-0.316051	nan	-0.200855	nan	nan	nan	nan	0.765834	nan	-0.060499	nan	-0.338792	-0.474869	-0.312395	-0.450562	0.631150	-0.184554	-0.496568	nan	1.025249	-0.191439	-0.393800	-0.014128	-0.392889	-0.322916	nan	-0.460293	-0.182935	-0.153168	-0.545048	-0.093180	nan	0.225497	nan	nan	-0.050547	-0.035623	nan	-0.429489	-0.651230	nan	0.109831	-0.253569	-0.304187	-0.757136	nan	-0.407953	-0.172880	0.225896	-0.124650	0.404438	-0.278338	0.676358	0.102604	nan	0.071175	0.701722	2.766895	-0.332718	0.200391	-0.113778	0.090591	-0.342221	0.440389	0.968416	0.152312	-0.438715	0.116453	-0.289726	nan	-0.127844	0.135441	-0.732415	nan	nan	0.369340	nan	nan	nan	nan	nan	-0.376654	-0.426711	nan	0.329170	0.088458	nan	-0.234051	0.203284	0.156843	nan	0.970065	nan	-0.098160	nan	0.014632	nan	0.256873	-0.049152	-0.260200	nan	nan	nan	0.626579	-0.285093	0.244340	0.201269	-0.342285	0.110783	0.158687	-0.453498	-0.354170	-0.111170	-0.062455	nan	0.170343	0.024254	-0.335108	0.156729	nan	0.557843	nan	1.352004	nan	-0.020119	-0.209953	-0.418717	0.107102	-0.168479	-0.042227	0.046235	0.253850	-0.084545	nan	1.178867	-0.103371	1.540236	-0.000352	-0.085181	nan	-0.485347	nan	-0.021827	-0.113681	-0.125274	-0.366496	-0.367804	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.119126	nan	-0.416085	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.005896	0.998665	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.217622	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.041767	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	0.163504	nan	nan	nan	nan	-0.486207	nan	nan	nan	nan	nan	0.865622	nan	nan	0.245497	-0.573961	nan	0.266594	nan	-0.025309	nan	0.024376	-0.275329	0.184680	-0.187755	-0.270358	0.595347	-0.121682	nan	0.906455	nan	nan	nan	-0.386199	0.166275	nan	-0.364857	-0.473669	nan	-0.082997	nan	nan	nan	1.106571	-0.391481	-0.357996	-0.298895	-0.150466	nan	0.157226	0.370765	-0.639713	-0.205941	0.170073	-0.233150	-0.005755	0.082972	0.155363	0.054915	0.130737	0.848404	-0.141295	-0.146291	-0.317983	0.005869	-0.225642	-0.027208	0.133381	-0.503229	0.500372	0.147946	0.223088	-0.227036	-0.085662	-0.484465	-0.187568	-0.192372	nan	-0.304203	nan	-0.543672	nan	nan	nan	-0.192604	-0.421335	-0.575204	nan	-0.451805	0.044478	nan	nan	-0.018559	0.154486	0.491860	nan	-0.262406	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.433638	nan	-0.316137	nan	-0.005388	-0.184509	-0.263857	nan	nan	nan	nan	nan	nan	-0.417204	nan	-0.467643	nan	nan	nan	-0.281332	0.048001	nan	0.074367	0.464403	-0.190363	nan	nan	-0.204196	-0.562790	-0.086740	nan	nan	nan	-0.441238	nan	0.003137	0.017722	-0.617849	-0.053200	-0.355265	0.305782	nan	0.382800	nan	nan	nan	nan	-0.316541	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	nan	-0.313869	nan	1.938720	-0.170564	nan	nan	-0.365692	-0.259854	-0.271620	nan	-0.033167	-0.425726	-0.337518	-0.508203	nan	-0.321046	nan	0.109863	-0.085295	nan	-0.392853	-0.332886	0.046866	nan	-0.072946	-0.282880	nan	0.437903	-0.390503	nan	-0.445866	nan	nan	-0.697174	-0.237883	0.996852	-0.021289	-0.249012	nan	-0.235864	-0.551922	-0.039140	0.132761	nan	-0.000585	-0.388119	nan	0.847094	nan	-0.440410	-0.267915	0.117292	nan	-0.729473	0.703680	nan	nan	-0.258475	0.516386	-0.510212	-0.103087	-0.213671	0.411249	0.717649	-0.228116	-0.192381	nan	-0.224747	-0.543443	-0.405085	nan	nan	nan	nan	nan	-0.246832	0.056778	-0.134523	nan	0.266096	nan	-0.053077	1.759381	nan	nan	0.512064	0.537340	-0.150449	nan	nan	nan	nan	nan	nan	-0.363980	-0.363345	nan	nan	0.234002	-0.166385	nan	nan	nan	nan	-0.539494	0.103373	-0.481606	nan	0.465333	0.099508	nan	nan	-0.502997	nan	-0.225889	0.493369	-0.622523	-0.385000	-0.153559	-0.134633	nan	-0.079860	-0.362093	-0.228914	-0.264698	0.105541	nan	-0.067154	-0.140849	-0.498545	0.202228	nan	0.739133	-0.509332	-0.263121	0.258930	nan	-0.679921	0.228373	-0.319420	-0.157416	-0.371906	-0.294867	-0.195848	nan	0.107341	-0.619119	nan	-0.179686	-0.236770	-0.564975	-0.423714	-0.194092	nan	nan	nan	nan	nan	-0.359516	-0.736095	-0.451625	nan	nan	nan	-0.508315	-0.452826	nan	-0.247064	nan	-0.496840	0.029668	-0.384303	-0.345193	-0.082610	-0.466420	nan	nan	-0.123249	-0.508771	-0.048365	-0.300055	nan	nan	-0.365758	nan	nan	nan	nan	nan	nan	nan	-0.209880	nan	nan	nan	nan	-0.305390	-0.462766	-0.509763	nan	-0.390991	nan	nan	-0.311137	nan	-0.105759	nan	-0.411670	-0.099623	-0.187737	0.181018	0.629400	nan	nan	nan	nan	-0.371103	nan	-0.285324	0.013751	-0.427397	-0.077225	-0.087638	0.645254	nan	nan	nan	nan	nan	nan	ACNNEWEDEQYEQYISFKSPIPAGGEGVTDIYVRYKEDGKVTYRLPIVVSGSTTNTLDREVHIAVDKDTLETLNIERFSIHRPGLWYTALEEDKYDFPKTVHISAGSSVEQLNIDFNLQGIDMLEKWVLPLTIVDDPSYNYKSHPRKNYAKALLRVIPFNNYSGSYTSSAMKVYTYINGKPDTNARTANKRTGYVINDNTVFFYAGLISEDLDKEVREKYKINVHFNEDGTLHMEQGDPTNKMKFELIGTPTYSKTSVMDATRPYLERRYVQIMFEYDFEDFSYGGNGVDVIPIKYKVEGSMTLQRNINTQIPDEDQQIEW	SP15308A-bot-339-19-339

As you can see, this dataset contains numerical values (Z-scores) corresponding to the ability of different lectins to bind different glycans. From these data, we can compute a heatmap with make_heatmap. The results should teach us whether some specific strains bind specific glycans. As we work with quantitative values, the datatype parameter must be set to 'response'. If you want to include literature-annotated motifs (e.g., VIM, Lewis X, Blood group A), just add ‘known’ to the list of the feature_set. Let’s see what kind of heatmap we can generate using this dataset.

make_heatmap(glycan_binding.iloc[:,:-1], mode = 'motif', feature_set = ['exhaustive'], datatype = 'response', yticklabels = 1,
            xticklabels = False)

Due to the high number of lectins in this dataset, the heatmap is difficult to analyze in detail. However, we can clearly see that there are clusters of lectins recognizing specific glycan motifs. Examples include Neu5Ac(a2-6)Gal, Neu5Ac(a2-3)Gal, Fuc(a1-6)GlcNAc, and others. All these represent well-known glycan motifs with important roles in biology.

Another way to analyze the glycan binding capacities of lectins is to use the get_pvals_motifs function from the glycowork.motif.analysis module. As we are working with data obtained from different proteins, we must also set multiple_samples to True.

results = get_pvals_motifs(glycan_binding.iloc[:,:-1], feature_set = ['exhaustive'], multiple_samples = True)

	motif	pval	corr_pval
58	b1-4	1.8105766142037072e-143	0.0
43	Neu5Ac	5.2714117926304566e-123	0.0
26	GlcNAc	1.980368513762096e-113	0.0
38	Man	2.8616948168475176e-107	0.0
205	Man(a1-3)Man	1.5247041217922468e-97	0.0
207	Man(a1-6)Man	8.114125848901856e-95	0.0
8	Gal	3.710533247731733e-91	0.0
215	Neu5Ac(a2-3)Gal	7.63292387136606e-85	0.0
183	GlcNAc(b1-4)GlcNAc	2.1665838781174787e-84	0.0
51	a1-6	2.6478638411565924e-79	0.0
208	Man(b1-4)GlcNAc	4.903488727619156e-76	0.0
109	Gal(b1-4)GlcNAc	1.6674581628358519e-75	0.0
56	b1-2	5.941018178401668e-72	0.0
219	Neu5Ac(a2-6)Gal	2.9660362110348526e-71	0.0
53	a2-6	3.8643923787825105e-65	0.0
52	a2-3	3.014034253371599e-60	0.0
173	GlcNAc(b1-2)Man	3.372781726327191e-60	0.0
175	GlcNAc(b1-3)Gal	4.7929799268331535e-58	0.0
57	b1-3	3.3478670790694135e-55	0.0
48	a1-3	3.8297399645018434e-36	0.0
188	GlcNAc(b1-6)Man	3.6666408458452625e-27	0.0
203	Man(a1-2)Man	1.3167886060718853e-25	0.0
111	Gal(b1-4)GlcNAc6S	1.7127576986244706e-20	0.0
83	Fuc(a1-6)GlcNAc	7.357723453995614e-19	0.0
204	Man(a1-3)GalNAc	1.432451207058872e-16	2.4091839634365897e-14

These results indicate, for each motif, a p-value and a corrected p-value. These quantitative results support our qualitative observations from before. However, this represents a bird’s eye view across all lectins in the dataset. A follow-up analysis might be to focus on a specific group of lectins, analyzing their binding patterns in depth.

Example 3: Glycan classification using machine learning models

The investigation of glycans from animals and viruses demonstrated how specific their presence in an organism can be. This particular characteristic can be exploited to predict the source of a given glycan among organisms. By classifying which glycans stem from which species/taxonomic groups, machine learning models can be trained to learn which motifs are enriched in which species, allowing for further insights.

Let’s see how we can use glycowork to classify vertebrate versus invertebrate glycans! First, we need to annotate each known glycan using a binary code where 1 means vertebrate glycan and 0 means invertebrate glycan.

vertebrate = [1 if k == 'Chordata' else 0 for k in df_species[df_species.Kingdom == 'Animalia'].Phylum.values.tolist()]

Now, we have to generate two sets of glycans that we will use as train set and test set. In addition, we also need their corresponding sets with classes labelled as 0 or 1. One way to do it is to use the general_split function from the glycowork.ml.train_test_split module. This function assigns the data with a 80/20 ratio to train and test sets respectively.

X_train, X_test, y_train, y_test = general_split(df_species[df_species.Kingdom == 'Animalia'].target.values.tolist(), vertebrate)

It is now time to train a machine learning model on these data! Different machine learning algorithms are available in glycowork and here we can use the train_ml_model function from the glycowork.ml.model_training module. This classification method uses a standard machine learning model (XGBoost) trained on short glycan motifs which may not be the most accurate, but can be sufficient for a preliminary exploration of our problem. Plus, it has the advantage of running faster than other more complex algorithms such as deep learning so… let’s give it a try!

model_ft, _, X_test = train_ml_model(X_train, X_test, y_train, y_test, feature_calc = True, feature_set = ['exhaustive'],
                         return_features = True)

Calculating Glycan Features...

Training model...
[16:09:22] WARNING: C:/Users/Administrator/workspace/xgboost-win64_release_1.4.0/src/learner.cc:1095: Starting in XGBoost 1.3.0, the default evaluation metric used with the objective 'binary:logistic' was changed from 'error' to 'logloss'. Explicitly set eval_metric if you'd like to restore the old behavior.

Evaluating model...
Accuracy of trained model on separate validation set: 0.927536231884058

As you can see, even this (quite) simple algorithm is enough to reach around 93% prediction accuracy on the test set! It also confirms that glycan sequence is highly representative of their origin and allows to efficiently discriminate vertebrate from invertebrate glycans.

One more step we can do is the analysis of the model we have generated. This is important, as we do not know yet what information is the most useful for the model to predict the origin of a glycan. The analyze_ml_model function can help us to understand on which basis the model predictions are made.

analyze_ml_model(model_ft)

Interesting! It appears that the presence/absence of the Gal(b1-4)GlcNAc motif (type II LacNAc) and GlcA in glycans is quite discriminative of its belonging to the Chordata phylum or not!

Next to knowing what is important for model prediction, it is often also crucial to know the limits of a model. In glycowork, you can use get_mismatch from the glycowork.ml.model_training module to get a sense of wrong classifications. This could for instance inform you about potential model biases.

get_mismatch(model_ft, X_test, y_test)

[('Man(a1-?)Man(a1-?)[Man(a1-?)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc',
  0.7171406745910645),
 ('Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc',
  0.919142484664917),
 ('Man(a1-?)Man(a1-?)[Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc',
  0.8076043128967285),
 ('Man(a1-?)Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc', 0.49124422669410706),
 ('GlcNAc(b1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc',
  0.46067529916763306),
 ('Gal(a1-4)Gal(b1-4)GlcNAc(b1-6)[Gal(b1-3)]GalNAc', 0.9600384831428528),
 ('Gal6Aep(a1-2)[Gal3Me(a1-3)]Gal(b1-4)Glc1Cer', 0.5451778769493103),
 ('Man(a1-2)Man(a1-6)Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc',
  0.4874219596385956),
 ('GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]Man',
  0.4899427592754364),
 ('Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)[Man(a1-2)Man(a1-3)]Man(b1-4)GlcNAc(b1-4)GlcNAc',
  0.591519832611084)]

Here we can see that our model was especially confused by glycans which can occur in both phyla and therefore are ambiguous for classification ambiguous. Maybe in these scenarios a more powerful, deep learning-based model might be able to differentiate (at least a bit better)!

Constructing and exploring biosynthetic networks

The network module of glycowork is designed to readily construct and analyze biosynthetic networks from a list of glycans in a very modular way. Below, this is illustrated with a fictional set of milk glycans (all default parameters of construct_network are optimized for milk glycans and should be changed for other glycan classes). Just inputting a list of biosynthetically related glycans into construct_network and then plot_network will build and plot our biosynthetic network! (depending on your browser etc, this might not render correctly)

glycans = ["Gal(b1-4)Glc-ol", "GlcNAc(b1-3)Gal(b1-4)Glc-ol",
           "GlcNAc6S(b1-3)Gal(b1-4)Glc-ol",
           "Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc-ol", "Fuc(a1-2)Gal(b1-4)Glc-ol",
          "Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)[Gal(b1-3)GlcNAc(b1-6)]Gal(b1-4)Glc-ol"]
network = construct_network(glycans)
plot_network(network, plot_format = 'kamada_kawai')

We can see that, in these networks, every dot is a glycan (and can be hovered over to see its sequence) and every link is the addition of a monosaccharide in a defined linkage by an enzyme. Blue dots are glycans which we provided as input, orange glycans are inferred in order to connect the network.

One thing you can immediately see is that there often are multiple paths to reach a blue dot, via several orange routes. This is because we don’t know the order in which the monosaccharides have been added. We show in recent work that this order is highly conserved. Therefore, we could use knowledge gathered over decades in the breast milk of over a hundred mammalian species, to find out which order is realistic.

glycowork can help you here as well, since it contains pre-computed biosynthetic networks for milk glycans of >100 mammalian species. We can use the evoprune_network function to use this information to prune our networks, removing paths that are evolutionarily very unlikely.

print(len(network.edges()))
network = evoprune_network(network)
print(len(network.edges()))
plot_network(network, plot_format = 'kamada_kawai')

19
11

Nice, that’s a bit more organized! evoprune_network has removed a few of the nodes that had zero probability. You can see that the orange nodes have changed size as well. This is to indicate their probability. If you change the threshold parameter of evoprune_network these low-probability nodes might also be pruned away, yet network connectivity is always preserved.

Now, let’s say we want to highlight parts of the network containing the monosaccharide Neu5Ac. For this, we can use highlight_network. With this function, you can highlight many things in a network: motifs, abundances, species occurence, evolutionary conservation, etc.

network = highlight_network(network, 'motif', motif = 'Neu5Ac')
plot_network(network, plot_format = 'kamada_kawai')

You can see that the node color has changed now. Every glycan/node that contains Neu5Ac now is showing up as a green node, while those lacking Neu5Ac are purple. You can also still make out which nodes are inferred (transparent) and actually observed (solid). The probabilities from evoprune_networkare also still contained within the network. Apart from these visualizations, you can use these networks for all kinds of analytical work, from motif.graph.generate_graph_features to network.evolution.get_communities; your only limitation is your creativity!:-)

We can also use the functions within glycowork.network to compare species and their milk glycomes: Let’s have a look at the two pre-computed biosynthetic networks (found in net_dic) of black bears (Ursus americanus) and brown bears (Ursus arctos).

blackbear_network = net_dic['Ursus_americanus']
brownbear_network = net_dic['Ursus_arctos']
plot_network(network = brownbear_network)

pydot2 threw an error (maybe you're using Windows?); we'll use kamada_kawai instead

As in our example we have two networks, it is possible to directly combine and compare them to identify conserved and specific nodes, using for instance network_alignment.

combined_network = network_alignment(network_a = blackbear_network, network_b = brownbear_network)
plot_network(network = combined_network)

pydot2 threw an error (maybe you're using Windows?); we'll use kamada_kawai instead

In this new representation, the blue nodes are specific to network_a (here the black bear), and orange nodes are specific to network_b (the brown bear). The remaining brown nodes are conserved in both species. Virtual nodes follow the same color scheme but are transparent. Here, we can see many blue nodes that are specific to the black bear. This is not so surprising, as the black bear network is larger.

We can also make use of the network.evolution module to compare the similarity of related species, such as different kinds of bears. Using the metric of Jaccard distance between their networks, we can retrieve sequences and networks for various bear species within glycowork with the function distance_from_metric. Then, we can use the distance matrix in the dendrogram_from_distance function to plot the resulting clustering. One immediate result here is that polar bears (Ursus maritimus) seem to have a very different milk glycome from other types of bears!

bears = [net_dic[k] for k in list(set(list(net_dic.keys()))) if 'Ursus' in k]
df_bears = df_species[df_species.Genus=='Ursus'].reset_index(drop = True)
dm = distance_from_metric(df_bears, bears, cut_off = 5)
dendrogram_from_distance(dm)

	Ursus_americanus	Ursus_arctos	Ursus_arctos_lasiotus	Ursus_arctos_yesoensis	Ursus_maritimus	Ursus_thibetanus_japonicus
Ursus_americanus	0.000000	0.647482	0.582677	0.613636	0.952000	0.647482
Ursus_arctos	0.647482	0.000000	0.422222	0.728070	0.955056	0.000000
Ursus_arctos_lasiotus	0.582677	0.422222	0.000000	0.731481	0.910256	0.422222
Ursus_arctos_yesoensis	0.613636	0.728070	0.731481	0.000000	0.777778	0.728070
Ursus_maritimus	0.952000	0.955056	0.910256	0.777778	0.000000	0.955056
Ursus_thibetanus_japonicus	0.647482	0.000000	0.422222	0.728070	0.955056	0.000000

Deep Learning Code Snippets

For those of you that are blessed with GPU-access, we wrote up a minimal example for training a glycan-focused deep learning model. Even if you usually do not have access to a GPU, you can always go to Google Colab (click on the badge at the top of this site to open this notebook on Google Colab) and change the runtime type to GPU. Then you can paste the following code in a cell, execute, and see our model train!

First, we need to install torch_geometric for deep learning (keep the package versions; especially if you want to use our trained models):

!pip install torch-scatter -f https://pytorch-geometric.com/whl/torch-1.13.0+cu116.html
!pip install torch-sparse -f https://pytorch-geometric.com/whl/torch-1.13.0+cu116.html
!pip install torch-cluster -f https://pytorch-geometric.com/whl/torch-1.13.0+cu116.html
!pip install torch-spline-conv -f https://pytorch-geometric.com/whl/torch-1.13.0+cu116.html
!pip install torch-geometric==2.2.0

SweetNet-type model

Then, we import the relevant glycowork modules:

from glycowork.glycan_data.loader import df_species
from glycowork.ml.train_test_split import hierarchy_filter
from glycowork.ml.processing import split_data_to_train
from glycowork.ml import models
from glycowork.ml import model_training

Finally, we can prepare the data and train a model to classify from which taxonomic kingdom a glycan came from, using our graph convolutional neural network, SweetNet.

train_x, val_x, train_y, val_y, id_val, class_list, class_converter = hierarchy_filter(df_species,
                                                                                       rank = 'Kingdom')

dataloaders = split_data_to_train(train_x, val_x, train_y, val_y)
model = models.prep_model('SweetNet', len(class_list))
optimizer_ft, scheduler, criterion = model_training.training_setup(model, 0.0005)
model_ft = model_training.train_model(model, dataloaders, criterion, optimizer_ft, scheduler,
                   num_epochs = 100)

And that’s it! You’ve just trained a state-of-the-art deep learning model with effectively five lines of code (excluding package imports of course).

LectinOracle-type model

For LectinOracle, you need a protein sequence and glycan sequences. Further, you will need to retrieve the ESM1b-representation for your protein sequence, as that is used as the actual input for LectinOracle. If you don’t want to do that, check below for the LectinOracle_flex model. Here the minimal process is shown for the arbitrary protein sequence “QWERTFVCF”. The trained model is yet again retrieved via prep_model.

!pip install fair-esm
import esm
from glycowork.ml.inference import get_esm1b_representations, get_lectin_preds
from glycowork.ml.models import prep_model

model, alphabet = esm.pretrained.esm1b_t33_650M_UR50S()
rep = get_esm1b_representations(["QWERTFVCF"], model, alphabet)
leor = prep_model("LectinOracle", 1, trained = True)
get_lectin_preds("QWERTFVCF", ['Neu5Ac(a2-6)GalNAc', 'Gal(b1-4)Glc'], leor, rep)

The output here (same as for LectinOracle_flex below) is a table/dataframe, where each row shows the tested glycan and its associated binding prediction. The higher the binding prediction, the more likely an interaction (negative values in general likely mean “no interaction”), with a moderate correlation with binding affinity (based on our findings so far).

LectinOracle_flex-type model

While it has a slightly lower performance than LectinOracle, LectinOracle_flex can directly use protein sequences as input (no ESM1b-conversion necessary). This is made possible by knowledge distillation, in which we train a sub-module to mimic the conversion of the much larger ESM1b model. Just get the correct model via prep_model and declare flex = True and you’re good to go!

from glycowork.ml.inference import get_lectin_preds
from glycowork.ml.models import prep_model
leor = prep_model("LectinOracle_flex", 1, trained = True)
get_lectin_preds("QWERTFVCF", ['Neu5Ac(a2-6)GalNAc', 'Gal(b1-4)Glc'], leor, flex = True)